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To browse Academia. Skip to main content. Log In Sign Up. Download Free PDF. Kinship and association in social foraging northwestern crows Corvus caurinus. Renee Ha. Download PDF. A short summary of this paper. Bird Behavior, Vol. We investigated the role of kinship and social affiliation on these find a dissertation on proquest by combining microsatellite DNA-based measures of relatedness with nicole kaib dissertation animal behavioral sam- pling of northwestern crows in the field.

Scrounges were either passive nicole kaib dissertation ground approach without pursuit or aggressive noisy approach with pursuit. The combination of individually marked animals and information on relatedness allowed investigation nicole kaib dissertation the roles of dominance and kinship in scrounging, including the target and nicole kaib dissertation of the scrounge. Crows in descriptive essay on family reunion groups were more likely to be associated i.

There was no difference between the average association of the forag- ing group and the average association between the scrounge interactors. There was also no difference in the average association between passive and aggressive scrounge interactions. Adults were more likely than juveniles writing up dissertation discussion demonstrate a preference cheap best essay ghostwriter for hire ca social companions, and males were more likely than females to demonstrate a preference for social companions.

Within forag- ing groups, passive scrounges tended to occur between more closely related individuals, and aggressive scrounges among less closely related individuals. Thus, scroungers modulate their mode of qualitative research dissertation chapters according to relatedness, or producers modulate their stephen crane writing style, or both.

They discriminate close from more distant relatives or unrelated individuals by adjusting the level of aggressiveness associated with theft. The pro- Giraldeau, Small prey items are quickly Hockey, In- nicole kaib dissertation distant locations converge on this beach dur- deed, this has rarely been addressed Sklepkovych, ing times of nicole kaib dissertation exposure. However, Hatch and Lefebvre ritorial, defending an area around prime nest sites. During the aggressive encounters, in- tion from kin. Sklepkovych found that adult dividuals physically attack one another and engage Siberian jays discriminated between kin and nonkin in chasing and nicole kaib dissertation. During a passive while foraging, specifically by tolerating juvenile scrounge, an individual simply displaces another kin over juvenile nonkin at bait sites.

We question the degree to which feeding sites, particularly related juveniles. Alter- our data reflect differential behavior towards rela- nicole kaib dissertation, being scrounged upon may be an nicole kaib dissertation tives or simply differential behavior towards those able cost to social foraging, and scroungers may nicole kaib dissertation with which scroungers are most familiar. This is sup- Thus, we have measured association nicole kaib dissertation groups ported by the vast literature on interspecific of animals in our population.

Thus, there are the degree writing up dissertation discussion affiliation, and thus familiarity, of vari- dissertationen online tbingen explanations for each hypothesis. If ous groups of animals. Indeed, this index provides scrounge interactions occur among unrelated pairwise measures of association parallel to nicole kaib dissertation interactors, then that would suggest that scrounging pairwise measures of kinship calculated from the is potentially a cost to social foraging but see DNA marker data.

Alternatively, if scrounge interac- lowed investigation of the roles of dominance, af- tions occur among related interactors, then this would filiation, nicole kaib dissertation kinship in scrounging, including the be more consistent with a kin selection argument target and form of the scrounge. Demonstra- egies in northwestern crows nicole kaib dissertation combining tion that indirect fitness effects may play a role in microsatellite DNA-based measures of nicole kaib dissertation social foraging nicole kaib dissertation would suggest that current with focal animal behavioral sampling in the field. WM; United States Department of the be the most accurate. We briefly restrained the animals gested that this index tends to nicole kaib dissertation associa- approximately 10 min writing the essay science banding and blood draw.

According to nicole kaib dissertation taken from the wing vein of each bird. Cairns and Schwager legs. Captured birds were held in an animal carrier suggested that the twice-weight index is the least and kept warm until they were processed nicole kaib dissertation had biased when sampling favors the sighting of indi- recovered from the sedation if applicableat which viduals A and B together. Finally, the simple index time they were released at the location at which they of association was considered by Ginsberg and had been captured. Young to be the least biased of all the indi- ces. They suggested that because it does not double Sexing the number of sightings in which both individuals Crows are not sexually dimorphic; thus, we de- are seen separately i.

The technique was vali- of the indices. Generic amplification reactions consisted of ng where x is the number of times individual A and B of template DNA, 0. Tetranucleotide loci were run on and B nicole kaib dissertation seen separately. These tests randomly generated alternative plied Biosystems Inc. To test for acrylamide gel. The data were analyzed using ABI long-term companionship, we calculated individual Genescan software. With this dard deviations of the association indices would in- method, relatedness coefficients r range from —1. A negative coefficient indicates that the in- observed association data were significantly higher dividuals share nicole kaib dissertation alleles than expected given the than the randomly permuted data, then the null hy- background paper research format of sharing in the population.

Note pothesis of no preferred companions between sam- nicole kaib dissertation r is a measure of relative relatedness within the pling periods was rejected Gowans, The study was conducted from Analysis of nicole kaib dissertation patterns and social struc- February 14, to August 19, Hal Whitehead of nicole kaib dissertation h for a total of nicole kaib dissertation. All observations Dalhousie University Whitehead, Coefficient were conducted by R. The date nicole kaib dissertation tempera- of association COA indices were calculated for 28 ture were recorded at the start of each session.

In- crows sighted two or more times. This foraging away from the previous subject. During phase of the study was conducted nicole kaib dissertation October 11, each 5-min period, the age class of the subject nicole kaib dissertation through November 1, All observation vs. The date and temperature were scrounge by focal animal, loss of prey to another recorded nicole kaib dissertation the start nicole kaib dissertation each session. The observer the number of scrounge attempts made by the fo- performed a s nicole kaib dissertation sample noting the number of cal animal or top creative essay editor services for college of scrounge attempts made crows on the beach and the identity of any marked on the focal animal by other animals were recorded individuals.

Following the census scan, banking case study con- on a laptop computer Compaq Contura us- ducted a nicole kaib dissertation period of all-occurrences sampling ing modified EVENT version 3. Age class was determined by plumage the nicole kaib dissertation of the scrounge actor and recipient. A s cen- Capture of arcada thesis guide prey item could nicole kaib dissertation through pro- sus scan was repeated at the end of the min pe- ducing or scrounging.

Producing was defined as an riod. Scroung- Data Analysis ing was defined as taking a prey item from a pro- ducer. Passive scrounges were those in which no The variances were homogeneous, the distribu- physical contact such as pecking or chasing oc- personal essay def were normal, and recorded relatedness values curred between the interactors and the individual nicole kaib dissertation considered independent events. Levels of re- who lost the prey item to a scrounger made no at- latedness among marked birds within larger groups tempt to retrieve nicole kaib dissertation item. Unsuccessful passive of unmarked birds were nicole kaib dissertation with t-tests.

Each scrounges were situations in which an individual t-test analysis used a different null hypothesis and with a food item was approached nicole kaib dissertation another indi- underlying statistical population, and thus no vidual with no physical contact or chasing and the Bonferroni adjustments were required. Frequency approaching bird was nicole kaib dissertation at obtaining nicole kaib dissertation data were analyzed with chi-square tests. Because prey item. Begging was associated with wing flicks, several chi-square analyses were performed on the bowing, and specific vocalizations. Begging was not same underlying statistical population, an overall nicole kaib dissertation as a scrounge in this study.

The relatedness and social nicole kaib dissertation by the scrounger in an attempt nicole kaib dissertation obtain the prey nicole kaib dissertation values by age and sex class categories were item or by the producer who lost the prey item to analyzed with ANOVA. All comparisons were two- the scrounger. When nicole kaib dissertation animal flew to a new loca- tailed. Presence or absence of Results other foragers was recorded. Social Affiliation Patterns Focal sampling alternated with s scan samples of the study site. During scan samples, we recorded The observed association of individuals in the the number of birds on the beach.

Adults ing pair and the average background relatedness of nicole kaib dissertation two or more times nicole kaib dissertation related to one an- the associated foraging group. Individuals that scrounged ag- Kinship, Sex, Age, and Scrounging gressively did select scrounging targets that were less Numerous scrounge attempts were observed in related to them —0.

However, the mean related- ness of the scrounger to its foraging group did not differ between passive and aggressive scrounges. The nicole kaib dissertation relatedness of individuals involved in ag- gressive encounters —0. There was no significant more related 0. There was also the interactors was higher in a passive interaction no significant difference between the probability of 0. Finally, there was no relationship be- their age class juvenile vs. The level of association in foraging groups where There nicole kaib dissertation no significant effect of season breed- a scrounge or scrounge attempt occurred was sig- doctoral dissertation vs.

However, there feeding pwc case study interview example during the breeding season. However, there on the nicole kaib dissertation of association within the essay on my favourite journey There was no relationship with season breeding vs.

Thus, the two most likely explanations of scrounging—fledgling feeding or dominance— Discussion do not fully explain the patterns observed here. Ad- ditionally, passive and aggressive scrounges were The average relatedness between producer and equally successful. If fledgling feeding was driving scrounger interactors was similar to that of the over- the system, one would expect that passive scrounges characteristics of case study research pdf essay on plastics population, and groups of foraging best college essay service were would be more successful.

If dominance were driv- not more closely nicole kaib dissertation than the background relat- ing the system, one would expect that aggressive edness of the population. However, when consider- scrounges would be more successful. We found no ing mean foraging group relatedness, passive evidence to support either of those explanations.

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